Multiple sclerosis (MS) is an autoimmune and neurodegenerative disease resulting in demyelination of neurons, lack of signal transmission and reduced muscular mobility. Although classically viewed as a T cell driven disease, recent findings suggest that B cells play a significant role in disease development and pathology. To further explore this, B cell profiles in peripheral blood from 28 treatment-naive MS patients were assessed using flow cytometry and compared to 17 healthy controls. Using conventional and algorithm-based analysis we observed a trend towards increased abundance of memory B cells (MBC) in MS. Further analysis revealed a significant increase of IgA+ MBC as well as minor differences in other MBC phenotypes. By screening circulating B cells for markers associated with B cell function, we discovered a significantly decreased expression of the B cell activation factor receptor (BAFF-R) in MS patients compared to controls. BAFF-R expression was negatively associated with serum levels of its ligand, suggesting that downregulation may be due to increased levels of BAFF. In healthy controls, BAFF-R expression was associated with abundance of differentiated MBC but this was not observed in MS. In MS patients, decreased BAFF-R expression correlated with increased production of proinflammatory TNF following B cell stimulation. Together, these data suggest that BAFF signalling is dysregulated in MS and may lead to aberrant MBC cell homeostasis and proinflammatory B cell activation. Previous attempts to block BAFF signalling in MS patients unexpectedly increased disease severity, suggesting that BAFF signalling contributes to MS immunopathology but that mechanisms are incompletely understood.