Background: Multiple regulatory mechanisms have been identified that contribute to outcomes of allergen-specific immunotherapy. Allergen-induced immune responses are thought to involve thousands of genes functioning within complex networks, and how these operate/integrate to maintain immunological homeostasis is incompletely understood.
Aim: To identify allergen-specific immunotherapy induced changes to Th-memory responses employing systems-level gene co-expression network analysis.
Methods: Genome-wide allergen-induced Th-cell responses were profiled prospectively during 24 months subcutaneous immunotherapy (SCIT) in 25 rhinitis. Molecular profiles of Th-cells were interrogated employing upstream regulator, pathways and co-expression network analysis.
Results: Prior to immunotherapy, mite-induced Th-cell response networks consisted of multiple discrete co-expression networks; Type1-IFN-, Th2-, Inflammation-, and FOXP3/IL2-associated signalling. A 109 gene signature correlated with symptom scores relating to cytokine signalling/T-cell activation-associated pathways. Upstream drivers demonstrated activation of key Th1/Th2- and Inflammation-associated genes. At 3.5mths SCIT updosing, symptoms reduced by 32.5% and minimal changes to pathway/upstream regulator profiles were detected. In contrast, network changes were evident, displaying merging of FOXP3/IL2- with Inflammation- and Th2-associated modules. By 12mths SCIT, symptoms had reduced by 41%, whilst pathway/upstream regulator/network profiles remained unchanged. At 24mths of SCIT, symptoms stabilised at 47% of baseline, and the Type1-IFN-associated network module merged into the Th2/FOXP3/IL2/Inflammation module.
Conclusions: Stabilisation of the clinical effectiveness of SCIT underlies progressive rewiring of mite-induced Th-cell-associated Th2-, FOXP3/IL2-, Inflammation- networks and Type1-IFN-signalling subnetwork. Importantly, integration only occurs following 24mths of SCIT treatment. The recruitment of Th2-antagonist Type1-IFN signalling into a coordinated expression network may be central to stabilising the clinical effects of SCIT.